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The CD1d-LSF-Hy19.3 TCR structure provides the first insight into lipid antigen recognition by a non-invariant NKT TCR. Surprisingly, the Hy19.3 TCR binding mode combines features of both iNKT cells, as well as MHC-restricted T cells, resulting in a unique antigen recognition mechanism among αβ TCRs. On one hand, in the type II NKT ternary complex both TCR α and β chain contact the CD1d molecule with a diagonal footprint, typical of MHC-TCR interactions
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